College of Science China Agricultural University Beijing 100083 China;
Key Laboratory of Adaptation and Evolution of Plateau Biota Northwest Institute of Plateau Biology Chinese Academy of Sciences Xining 810008 China;
Institute of Systems Science Academy of Mathematics and Systems Science Chinese Academy of Sciences Beijing 100190 China;
College of Mathematics and Systems Science Xinjiang University Urumuchi 830046 China;
College of Science China Agricultural University Beijing 100083 China.;
