Key Laboratory of Aquatic Botany and Watershed Ecology Wuhan Botanical Garden Chinese Academy of Sciences Wuhan 430074 China;
Hubei Key Laboratory of Wetland Evolution & Ecological Restoration Wuhan Botanical Garden Chinese Academy of Sciences Wuhan 430074 China;
Donghu Experimental Station of Lake Ecosystems State Key Laboratory of Freshwater Ecology and Biotechnology Institute of Hydrobiology Chinese Academy of Sciences Wuhan 430072 China;
Institute of Hydroecology Ministry of Water Resources & Chinese Academy of Sciences Wuhan 430079 China;
Faculty of Agriculture Laboratory of Plant Conservation Science Meijo University Aichi 468-8502 Japan;