of;
Education;
Key;
Laboratory;
for;
Biodiversity;
Science;
and;
Ecological;
Engineering,;
School;
of;
Life;
Sciences,;
Fudan;
University,;
Shanghai;
200433,;
China;
of;
Life;
Sciences,;
Guangxi;
Normal;
University,;
Guilin;
541004,;
China;
of;
Biodiversity;
Science;
and;
Geobiology,;
Tibet;
University,;
Lhasa;
850000,;
China;
of;
Ecology;
and;
Geobotany,;
School;
of;
Life;
Sciences,;
Yunnan;
University,;
Kunming;
650091,;
China;