Institute;
of;
Applied;
Ecology,;
Chinese;
Academy;
ofSeiences,;
Shenyang;
110016,;
China;
Graduate;
University;
of;
the;
Chinese;
Academy;
of;
Sciences,;
Beijing;
100049,;
China;
Key;
Laboratory;
of;
Plant;
Resources;
Conservation;
and;
Sustainable;
Utilization,;
South;
China;
Botanical;
Garden,;
Chinese;
Academy;
of;
Sciences,;
Guangzhou;
510650,;
China;
College;
of;
Life;
&;
Environment;
Sciences,;
Shanghai;
Normal;
University,;
Shanghai;
200234,;
China;