Tianjin Key Laboratory on Technologies Enabling Development of Clinical Therapeutics and Diagnosis,School of Pharmacy, Tianjin Medical University;
Shanghai Institute of Materia Medica, Chinese Academy of Sciences;
College of Pharmacy, The University of Michigan;
College of Pharmacy and Research Institute of Pharmaceutical Sciences, Gyeongsang National University;
College of Pharmacy, Sunchon National University;
Intranasal protein delivery; Brain targeting; Cell-penetrating peptide; Low molecular weight protamine; Blood–brain barrier;