College of Life Sciences, Liaocheng University, Liaocheng 252059, China;
Key Laboratory of Ecology and Biodiversity of Shandong Colleges and University, Liaocheng 252059, China;
College of Life Sciences, Liaocheng University, Liaocheng 252059, China;
Key Laboratory of Ecology and Biodiversity of Shandong Colleges and University, Liaocheng 252059, China;